Author contributions
H.I. to and M.S. (Michito Shimozuru) conceived of the study. N.M., M.Y., M.N., Y.Y., T.T., M.S. (Michito Shimozuru) and S.N. were involved in sample collection. J.Y., H.I., M.I.-M., and H.Q. supported with the technical aspects of the experiment. S.N. performed all laboratory work and constructed each age estimation model. S.N. and M.S. (Michito Shimozuru) wrote the article with inputs from J.Y., T.T., M.S. (Mariko Sashika), H.I. and M.I.-M. All authors reviewed the article.