AUTHOR CONTRIBUTIONS
MICY, MG and IG conceptualized the project. MG and IG led the study. These authors acted as co-second authors: EMI, AB, NT, NSY, NG, MY, IE. ICY, designed protocols, and carried out experiments together with, BG, BS, TB, YA, IB, AGB, BI, HAS, BK, YC, TY, IA, ICA, and GA. Under the supervision of PK, SBC, ECD and MG conducted histopathological investigations and performed TEM analyses. OE, provided inactivated SARS-CoV-2 virus samples. FC, supervised VNT assays and carried out experiments together with SU, HE, CA. Challenge experiments were supervised by ST and carried out by HP and ISY. HEZ coordinated the GMP production of VLPs in Nobel Pharma. MG and IG wrote the manuscript, with input from all authors..