Acknowledgements
The authors wish to thank the staff of the Research Group in Zoology (GIZ), Universidad del Tolima (Ibagué, Colombia). LAR-F and LVG-H thank the Convocatoria No. 755, 2016 para la Formación de Capital Humano de Alto Nivel para el Departamento del Tolima, Colfuturo (miniciencias) and Departamento del Tolima for support (doctoral scholarships). We thank the biologists Emmanuel Quintero, David Santiago Varón, Carlos Guzmán and Benjamín Polanco for their support during the field activities and the people from each of the localities surveyed for their hospitality. Specimens collected are covered through the “Permiso Marco de Recolección” granted to the Universidad del Tolima (Resolución No. 2191. November 27, 2018. Autoridad Nacional de Licencias Ambientales—ANLA). We thank Dr Jairo Pérez for allowing access to the voucher specimens of the Museo Javeriano de Historia Natural ”Lorenzo Uribe, SJ”.