Data Accessibilty Statement: All data from this project will be stored on Dryad at the time of acceptance.
Competing Interests Statement: The authors declare no competing interests
Author Contributions: JRG and BPH conceived and designed the experiments. KEV processed all samples, recorded and processed relevant data, and performed preliminary analyses and interpretation. JRG performed final data analysis and wrote the manuscript. BPH provided editorial advice.
Acknowledgements: The authors would like to thanks Mr. Marcel Verleur at Sappi International for invaluable access and support. This work was supported by the Tree Protection Cooperative Program at the Forestry and Agricultural Biotechnology Institute (FABI), University of Pretoria, South Africa.