Data Accessibilty Statement: All data from this project will be
stored on Dryad at the time of acceptance.
Competing Interests Statement: The authors declare no competing
interests
Author Contributions: JRG and BPH conceived and designed the
experiments. KEV processed all samples, recorded and processed relevant
data, and performed preliminary analyses and interpretation. JRG
performed final data analysis and wrote the manuscript. BPH provided
editorial advice.
Acknowledgements: The authors would like to thanks Mr. Marcel
Verleur at Sappi International for invaluable access and support. This
work was supported by the Tree Protection Cooperative Program at the
Forestry and Agricultural Biotechnology Institute (FABI), University of
Pretoria, South Africa.