ACKNOWLEDGEMENTS

We thank Da-Yong Zhang, Wayne Dawson, Yanjie Liu, Nianxun Xi, Oliver Bossdorf, and Wenjing Hong for their comments on the manuscript. We also thank the reviewers for the constructive suggestions in the revision. This study is supported by the National Natural Science Foundation of China (31600291) and the Fundamental Research Funds for the Central Universities in China (20720170074).